Tomb III, with two vaulted chambers, was the resting place of a young teenager, most likely a male.
Both of the larger tombs contained gold, silver, and ivory ornaments, as well as ceramic and metal vessels.
"[Andronikos] presented his theories [that the tombs were those of Alexander's father and his family] with great skill, and the Greek nation responded with fervent enthusiasm," Borza said.
"Indeed I was one of those who, in two early articles in the late 1970s, accepted Andronikos' view that the remains were those of Philip II."
Borza started to doubt Andronikos' conclusions, however, as he studied the evidence.
He contacted Olga Palagia, an art historian at the University of Athens, to evaluate the tombs' construction, pottery, and paintings.
Soon the duo realized the significance of the fact that Tomb II and Tomb III were built using a curved ceilings called barrel vaults.
"The earliest securely dated barrel vault in Greece dates to the late 320s [B.C.], nearly a generation after the death of Philip II," Borza told National Geographic News.
Palagia also found that paintings on the exterior frieze of the tomb reflected themes that were likely from the age of Alexander the Great, rather than that of his father.
The paintings depict a ritual hunt scene with Asian themes, suggesting influences resulting from Alexander's extensive campaigns to the east.
(Read related story: "Alexander the Great Conquered City via Sunken Sandbar" [May 15, 2007].)
The six-foot (two-meter) scepter found at the burial site is another clue, Borza added.
"We have several surviving coins issued in his own lifetime showing Alexander holding what appears to be a scepter of about that height," he said.
Additionally, a number of silver vessels discovered in Tomb II and Tomb III are inscribed with their ancient weights, which use a measurement system introduced by Alexander the Great a generation after Philip II's death.
"Once we have determined on archaeological grounds that Tomb II is a generation later than Philip II's death, we can then ask, Whose tomb is it?" Borza said.
"We have a double royal burial from this era attested in the ancient literature. Thus the tomb is that of [Alexander's half brother] Philip III Arrhidaeus and his queen, Adea Eurydice."
Borza and Palagia discussed their new analysis at the meeting of the Archaeological Institute of America in January. Their findings will be published in a forthcoming study from the German Archaeological Institute.
Most of the ancient artifacts found at Vergina are on display today at a museum at the site of the tombs.
Death of Alexander
Alexander died of disease in ancient Babylon, near modern-day Baghdad, Iraq, in 323 B.C.
His generals appointed Philip III to take his place, and the half brother claimed Alexander's royal objects as public symbols to solidify his power, historians suggest.
Alexander's son, Alexander IV, who was appointed joint king along with Philip III, was assassinated around 310 B.C. He is likely buried in Vergina's Tomb III, which contains the remains of a young teenager, Borza said.
Historically, the only known Macedonian royal teenage burial is that of Alexander IV, he explained.
Alexander's father, Phillip II, is buried in Tomb I, along with his wife and their infant, according to Borza.
"Tomb I is from the age of Philip II—unlike the big chamber tombs, which are later—and the human remains of the three burials accord well with the assassinations of these individuals."
Winthrop Lindsay Adams, a professor of history at the University of Utah who was not involved with the study, said Borza's work builds on what other specialists have thought about the various aspects of the Vergina tombs.
The work of Borza and his colleagues convincingly make the case that Tomb II is the final resting place of Alexander's half brother, Adams explained.
"Indeed for most scholars working in fourth-century Macedonia, the original attribution by Andronikos now seems doubtful," he said. "This case is convincing."
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